It is correct that genetic segregation analysis in autotetraploids requires considering the possibility of multivalent formation during pachytene and a potential double reduction of genetic loci. Both special conditions of autotetraploid meiosis are mentioned in [1
], p. 1-2. We agree that a more detailed tetrasomic linkage analysis could improve the data analysis for segregation in autotetraploids. We lack the mathematical expertise to critically review the different theoretical models presented in [2
] and refer to a detailed comment in [3
]. However, the evaluation of meiotic recombination suggested by [2
] also results in significantly higher rates in auto- and allopolyploids on one side and diploids on the other. Therefore, the claim in [2
] for 'qualitative differences from the original analysis' is not justified.
Further, the statement that we 'concluded that meiotic recombination was more frequent in the allotetraploids than in the autotetraploids' is incorrect: we wrote that the increase of recombination frequency of both polyploids over diploids was 'in the same range' (p. 4), which holds true for both different evaluations [1
Third, the green marker on the tester chromosome is further (distal) from the centromere [1
], not 'nearer to the centromere than the red marker' as assumed in [2
We have corrected three values in Table 1 according to the comments in [3
In summary, we are confident that the conclusions from our work remain unchanged.